Dr. Maclyn McCarty, 93, a pioneer in genetic research who helped demonstrate that genes are composed of DNA, died Sunday in New York City.
With Dr. Oswald Avery and Dr. Colin MacLeod, he conducted experiments in the 1940s on pneumococcus that pinpointed DNA as the carrier of genetic information.
The team's work paved the way for studies a decade later by Dr. James Watson and Dr. Francis Crick that revealed DNA's double helix structure, and for countless genetic studies since.
Dr. McCarty won the $100,000 Wolf Prize in medicine in 1990. The award foundation said the work with his partners had "opened the way to the great discoveries of the 20th century in molecular biology, climaxed by the unraveling of the genetic code."
Dr. McCarty was a 1937 graduate of the Johns Hopkins School of Medicine.
Among his three surviving children is a son in Baltimore, Richard E. McCarty, a professor and former dean and biology department chairman at the Johns Hopkins Unversity. Another son here, former state Del. Maclyn McCarty Jr., died in 2002.