Maryland’s pre-spring depth chart listed Joe Vellano and Keith Bowers as the Terps’ starting defensive ends. Bowers, however, was ruled out of practice last month with heart palpitations. Now Vellano will join him on the sideline.

“Joe Vellano is gonna be out for the spring with a foot injury,” Maryland coach Randy Edsall said in a YouTube video posted Saturday. “So that was disappointing.”

Vellano, a senior from Rexford, N.Y., was a first-team All-ACC selection last fall at defensive end. He appeared in all 12 games for Maryland during the 2011 season, contributing 94 tackles (54 solo), including 7.5 tackles for loss and 2.5 sacks. Vellano also broke up four passes, recovered two fumbles and forced two fumbles.

In other injury news, sophomore linebacker Alex Twine returned to practice and scrimmaged Saturday. Edsall said sophomore linebacker Cole Farrand “is getting a little bit closer” to returning, while senior wide receiver Kerry Boykins “took a shot to the head the other day, so he didn’t scrimmage” Saturday.

“Hopefully we’ll get some of those guys back and then stay healthy throughout the rest of the spring,” Edsall said.

At defensive end, juniors Isaiah Ross and Justin Anderson, and freshman Quinton Jefferson, were listed behind Vellano in Edsall’s initial spring depth chart. All three are coming back from injuries. Ross was injured in Maryland’s win over Miami last season and received a medical redshirt, while Anderson also missed the 2011 campaign.

Jefferson was originally a 2011 signee who deferred his enrollment while dealing with an undisclosed medical issue. The Pittsburgh native enrolled at College Park in January and has impressed Edsall thus far. Check out a video of Jefferson from spring practice below.