The afternoon heat was sweltering, so USC probably welcomed the chance to play a night game Saturday against Washington State.

But for Coach Lane Kiffin, there was no relief.

Not after Washington State defeated the No. 25 Trojans, 10-7, before a crowd that booed his play-calling throughout the game and chanted “Fire Kiffin” with just more than a minute left.

Athletic Director Pat Haden said before the season that the embattled Kiffin was not on the hot seat after last season's 7-6 finish. But fans were not shy about making their feelings known during a Pac-12 Conference opener that saw the Trojans fall to 1-1.

BOX SCORE: Washington State 10, USC 7

“You can’t worry about that,” Kiffin said of the boos. “It is what it is. I think heard those before the game started, in warmups actually. So I’m getting used to it.”

Washington State cornerback Damante Horton returned an interception for a touchdown late in the first half and sealed the victory with another after Andrew Furney kicked a field goal with just over three minutes left.

The defeat marked the Trojans' first loss to Washington State since 2002 and the first time they were held under seven points since a 10-6 loss to Utah in the 2001 Las Vegas Bowl.

USC’s 193 yards was its fewest since totaling 189 against Florida State in 1998. The Trojans’ 54 passing yards were their fewest since a paltry 23 in the same Florida State game.

“When your offense only passes for 50 yards, that’s just embarrassing,” quarterback Cody Kessler said.

Said quarterback Max Wittek: “It all comes down to execution. If we execute we don’t have to hear boos.”

Until USC figures out its quarterback situation and Kiffin believes the offensive line is capable of holding off defenders so the Trojans can get the ball downfield to receiver Marqise Lee, this might became a familiar scenario.

A defense that intercepted four passes in the season-opening victory over Hawaii intercepted two more Saturday and did not give up a touchdown.

The Trojans also pressured Washington State quarterback Connor Halliday into a fumble and followed up last week's seven-sack performance with four more.

A Washington State offense that generated 464 yards last week against Auburn was held to 222 by the Trojans, just seven rushing.

But Lee, the Biletnikoff Award winner, caught seven passes for only 27 yards against a defense that dropped eight players into coverage.

“We’re trying to get the passing game situated before we try to go vertical,” Lee said. “Try to get the little things before we try to go for the big things.”

Tailback Tre Madden was about the only offensive bright spot, the sophomore following up his 109-yard effort against Hawaii with 151 yards in 32 carries against Washington State. Several of Madden's gains came out of the wildcat formation.

Madden, however, was in no mood to celebrate.